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摩尔庄园的天空树屋里，小鼹鼠摩乐乐和养父菩提大伯过着平静的生活。由于一次误会，摩乐乐错失了出国旅行的好机会，负气伤心的乐乐，突然收到一张看似意外的火车票。接着，他召唤来了一辆从天而降的列车。这趟神秘的列车究竟有着怎样的故事呢？摩乐乐和小伙伴们的探险上天入地，还遭遇凶悍空匪的半路拦截，而列车最终的目的地又在哪里？麦田怪圈的问号、沙漠小镇的奇遇、虫洞飞跃的惊奇……。The film chronicles George's adventures as he befriends Kayla, a baby elephant, at a magic circus show and helps her travel across the country to be reunited with her family. Accompanied by his friend, the Man with the Yellow Hat, George travels by foot, train, and truck to reach Kayla's brother and sister in California, only to be accused of elephant-napping and brought all th...。