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在一劳永逸地击败了逆闪后，闪电侠巴里·艾伦(格兰特·古斯汀 Grant Gustin 饰)和艾瑞斯(坎迪斯·帕顿 Candice Patton 饰)的关系也比以往任何时候都要更加亲密。但是，当一群致命的无赖帮成员在一位名为“红死魔”的全新威胁的带领下袭击中心城时，巴里和他的闪电小队必须再次克服万难以便拯救中心城。 而由于此前消灭了逆闪导致负神速力失去了自己的代理人，一个未知的新对手也即将崛起，他的出现也将会是巴里和他的闪电小队迄今所面临的最艰苦的战斗，他们又能否再次拯救中心城呢？。While rowing out in the ocean, Mitch rescues an underwater photographer, named Linda, floating on some wreckage. Linda's somewhat disjointed, frantic story about something attacking her in an underwater cave which killed her partner and destroyed her boat is misconstructed by Kay Morgan, an ambitious, publicity seeking newspaper reporter, who creates a media frenzy by starting rumors about a "creature in the bay." Harvey Miller, the new lifeguard and prankster, benefits from the events by selling undersea creature T-shirts. Meanwhile, Shauni rescues a young girl from drowning and soon becomes compelled to rescue the girl, her brother and their mother from the life they live in the slums of South-Central L.A., much to Eddie's chagrin. While Shauni deals with her possessive father who disapproves of her lifeguard job, Mitch also pressures Captain Thorp to ask for funding for new advanced motorboats, called Scarabs, for the team.。